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Robert Lloyd
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35
Sharples, Gary
13
McGlynn, Peter
10
Rafferty, John
10
Bolt, Edward
9
Benson, Fiona
8
Sedelnikova, Svetlana
8
Mahdi, Akeel
6
Ingleston, Stuart
6
Rice, David
6
Moore, Timothy
5
Briggs, Geoffrey
5
Hargreaves, David
5
West, Stephen
4
Wen, Qin
4
Vincent, Simon
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All Publications
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2009: Rudolph Christian J; Upton Amy L; Harris Lynda; Lloyd Robert G
Pathological replication in cells lacking RecG DNA translocase.
Molecular microbiology 2009;73(3):352-66.
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2008: Rudolph Christian J; Upton Amy L; Lloyd Robert G
Maintaining replication fork integrity in UV-irradiated Escherichia coli cells.
DNA repair 2008;7(9):1589-602.
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2008: Grove Jane I; Harris Lynda; Buckman Carol; Lloyd Robert G
DNA double strand break repair and crossing over mediated by RuvABC resolvase and RecG translocase.
DNA repair 2008;7(9):1517-30.
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2007: Rudolph Christian J; Dhillon Paraminder; Moore Timothy; Lloyd Robert G
Avoiding and resolving conflicts between DNA replication and transcription.
DNA repair 2007;6(7):981-93.
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2007: Briggs Geoffrey S; McEwan Paul A; Yu Jing; Moore Timothy; Emsley Jonas; Lloyd Robert G
Ring structure of the Escherichia coli DNA-binding protein RdgC associated with recombination and replication fork repair.
The Journal of biological chemistry 2007;282(17):12353-7.
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2007: Rudolph Christian J; Upton Amy L; Lloyd Robert G
Replication fork stalling and cell cycle arrest in UV-irradiated Escherichia coli.
Genes & development 2007;21(6):668-81.
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2006: Mahdi Akeel A; Buckman Carol; Harris Lynda; Lloyd Robert G
Rep and PriA helicase activities prevent RecA from provoking unnecessary recombination during replication fork repair.
Genes & development 2006;20(15):2135-47.
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2006: Macmaster Rachel; Sedelnikova Svetlana; Baker Patrick J; Bolt Edward L; Lloyd Robert G; Rafferty John B
RusA Holliday junction resolvase: DNA complex structure--insights into selectivity and specificity.
Nucleic acids research 2006;34(19):5577-84.
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2005: Trautinger Brigitte W; Jaktaji Razieh P; Rusakova Ekaterina; Lloyd Robert G
RNA polymerase modulators and DNA repair activities resolve conflicts between DNA replication and transcription.
Molecular cell 2005;19(2):247-58.
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2005: Meddows Tom R; Savory Andrew P; Grove Jane I; Moore Timothy; Lloyd Robert G
RecN protein and transcription factor DksA combine to promote faithful recombinational repair of DNA double-strand breaks.
Molecular microbiology 2005;57(1):97-110.
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2005: Tessmer Ingrid; Moore Timothy; Lloyd Robert G; Wilson Anna; Erie Dorothy A; Allen Stephanie; Tendler Saul J B
AFM studies on the role of the protein RdgC in bacterial DNA recombination.
Journal of molecular biology 2005;350(2):254-62.
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2005: Briggs Geoffrey S; Mahdi Akeel A; Wen Qin; Lloyd Robert G
DNA binding by the substrate specificity (wedge) domain of RecG helicase suggests a role in processivity.
The Journal of biological chemistry 2005;280(14):13921-7.
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2005: Wen Qin; Mahdi Akeel A; Briggs Geoffrey S; Sharples Gary J; Lloyd Robert G
Conservation of RecG activity from pathogens to hyperthermophiles.
DNA repair 2005;4(1):23-31.
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2004: Meddows Tom R; Savory Andrew P; Lloyd Robert G
RecG helicase promotes DNA double-strand break repair.
Molecular microbiology 2004;52(1):119-32.
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2004: Allers Thorsten; Ngo Hien-Ping; Mevarech Moshe; Lloyd Robert G
Development of additional selectable markers for the halophilic archaeon Haloferax volcanii based on the leuB and trpA genes.
Applied and environmental microbiology 2004;70(2):943-53.
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2004: Moore Timothy; Sharples Gary J; Lloyd Robert G
DNA binding by the meningococcal RdgC protein, associated with pilin antigenic variation.
Journal of bacteriology 2004;186(3):870-4.
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2004: Briggs Geoffrey S; Mahdi Akeel A; Weller Geoffrey R; Wen Qin; Lloyd Robert G
Interplay between DNA replication, recombination and repair based on the structure of RecG helicase.
Philosophical transactions of the Royal Society of London. Series B, Biological sciences 2004;359(1441):49-59.
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2003: Rafferty John B; Bolt Edward L; Muranova Tatyana A; Sedelnikova Svetlana E; Leonard Philip; Pasquo Alessandra; Baker Patrick J; Rice David W; Sharples Gary J; Lloyd Robert G
The structure of Escherichia coli RusA endonuclease reveals a new Holliday junction DNA binding fold.
Structure (London, England : 1993) 2003;11(12):1557-67.
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2003: Muranova Tatyana A; Sedelnikova Svetlana E; Leonard Philip M; Pasquo Alessandra; Bolt Edward L; Lloyd Robert G; Rafferty John B
Crystallization of RusA Holliday junction resolvase from Escherichia coli.
Acta crystallographica. Section D, Biological crystallography 2003;59(Pt 12):2262-4.
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2003: Jaktaji Razieh P; Lloyd Robert G
PriA supports two distinct pathways for replication restart in UV-irradiated Escherichia coli cells.
Molecular microbiology 2003;47(4):1091-100.
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2003: Moore Timothy; McGlynn Peter; Ngo Hien-Ping; Sharples Gary J; Lloyd Robert G
The RdgC protein of Escherichia coli binds DNA and counters a toxic effect of RecFOR in strains lacking the replication restart protein PriA.
The EMBO journal 2003;22(3):735-45.
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2003: Mahdi Akeel A; Briggs Geoffrey S; Sharples Gary J; Wen Qin; Lloyd Robert G
A model for dsDNA translocation revealed by a structural motif common to RecG and Mfd proteins.
The EMBO journal 2003;22(3):724-34.
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2002: Trautinger Brigitte W; Lloyd Robert G
Modulation of DNA repair by mutations flanking the DNA channel through RNA polymerase.
The EMBO journal 2002;21(24):6944-53.
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2002: Dickman Mark J; Ingleston Stuart M; Sedelnikova Svetlana E; Rafferty John B; Lloyd Robert G; Grasby Jane A; Hornby David P
The RuvABC resolvasome.
European journal of biochemistry / FEBS 2002;269(22):5492-501.
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2002: McGlynn Peter; Lloyd Robert G
Recombinational repair and restart of damaged replication forks.
Nature reviews. Molecular cell biology 2002;3(11):859-70.
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2002: McGlynn Peter; Lloyd Robert G
Replicating past lesions in DNA.
Molecular cell 2002;10(4):700-1.
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2002: McGlynn Peter; Lloyd Robert G
Genome stability and the processing of damaged replication forks by RecG.
Trends in genetics : TIG 2002;18(8):413-9.
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2002: Bolt Edward L; Lloyd Robert G
Substrate specificity of RusA resolvase reveals the DNA structures targeted by RuvAB and RecG in vivo.
Molecular cell 2002;10(1):187-98.
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2002: Sharples Gary J; Bolt Edward L; Lloyd Robert G
RusA proteins from the extreme thermophile Aquifex aeolicus and lactococcal phage r1t resolve Holliday junctions.
Molecular microbiology 2002;44(2):549-59.
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2002: Ingleston Stuart M; Dickman Mark J; Grasby Jane A; Hornby David P; Sharples Gary J; Lloyd Robert G
Holliday junction binding and processing by the RuvA protein of Mycoplasma pneumoniae.
European journal of biochemistry / FEBS 2002;269(5):1525-33.
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2002: Gregg Amanda V; McGlynn Peter; Jaktaji Razieh P; Lloyd Robert G
Direct rescue of stalled DNA replication forks via the combined action of PriA and RecG helicase activities.
Molecular cell 2002;9(2):241-51.
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2001: McGlynn P; Lloyd R G
Action of RuvAB at replication fork structures.
The Journal of biological chemistry 2001;276(45):41938-44.
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2001: McGlynn P; Lloyd R G; Marians K J
Formation of Holliday junctions by regression of nascent DNA in intermediates containing stalled replication forks: RecG stimulates regression even when the DNA is negatively supercoiled.
Proceedings of the National Academy of Sciences of the United States of America 2001;98(15):8235-40.
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2001: McGlynn P; Lloyd R G
Rescue of stalled replication forks by RecG: simultaneous translocation on the leading and lagging strand templates supports an active DNA unwinding model of fork reversal and Holliday junction formation.
Proceedings of the National Academy of Sciences of the United States of America 2001;98(15):8227-34.
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2001: Bolt E L; Lloyd R G; Sharples G J
Genetic analysis of an archaeal Holliday junction resolvase in Escherichia coli.
Journal of molecular biology 2001;310(3):577-89.
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2000: Bolt E L; Sharples G J; Lloyd R G
Analysis of conserved basic residues associated with DNA binding (Arg69) and catalysis (Lys76) by the RusA holliday junction resolvase.
Journal of molecular biology 2000;304(2):165-76.
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2000: Ingleston S M; Sharples G J; Lloyd R G
The acidic pin of RuvA modulates Holliday junction binding and processing by the RuvABC resolvasome.
The EMBO journal 2000;19(22):6266-74.
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2000: McGlynn P; Mahdi A A; Lloyd R G
Characterisation of the catalytically active form of RecG helicase.
Nucleic acids research 2000;28(12):2324-32.
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2000: McGlynn P; Lloyd R G
Modulation of RNA polymerase by (p)ppGpp reveals a RecG-dependent mechanism for replication fork progression.
Cell 2000;101(1):35-45.
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1999: Sharples G J; Ingleston S M; Lloyd R G
Holliday junction processing in bacteria: insights from the evolutionary conservation of RuvABC, RecG, and RusA.
Journal of bacteriology 1999;181(18):5543-50.
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1999: McGlynn P; Lloyd R G
RecG helicase activity at three- and four-strand DNA structures.
Nucleic acids research 1999;27(15):3049-56.
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1999: Bolt E L; Sharples G J; Lloyd R G
Identification of three aspartic acid residues essential for catalysis by the RusA holliday junction resolvase.
Journal of molecular biology 1999;286(2):403-15.
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1999: Hargreaves D; Rafferty J B; Sedelnikova S E; Lloyd R G; Rice D W
Crystallization of Escherichia coli RuvA complexed with a synthetic Holliday junction.
Acta crystallographica. Section D, Biological crystallography 1999;55(Pt 1):263-5.
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1998: Hagan N F; Vincent S D; Ingleston S M; Sharples G J; Bennett R J; West S C; Lloyd R G
Sequence-specificity of Holliday junction resolution: identification of RuvC mutants defective in metal binding and target site recognition.
Journal of molecular biology 1998;281(1):17-29.
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1998: Whitby M C; Lloyd R G
Targeting Holliday junctions by the RecG branch migration protein of Escherichia coli.
The Journal of biological chemistry 1998;273(31):19729-39.
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1998: Hargreaves D; Rice D W; Sedelnikova S E; Artymiuk P J; Lloyd R G; Rafferty J B
Crystal structure of E.coli RuvA with bound DNA Holliday junction at 6 A resolution.
Nature structural biology 1998;5(6):441-6.
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1998: Rafferty J B; Ingleston S M; Hargreaves D; Artymiuk P J; Sharples G J; Lloyd R G; Rice D W
Structural similarities between Escherichia coli RuvA protein and other DNA-binding proteins and a mutational analysis of its binding to the holliday junction.
Journal of molecular biology 1998;278(1):105-16.
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1998: Chan S N; Vincent S D; Lloyd R G
Recognition and manipulation of branched DNA by the RusA Holliday junction resolvase of Escherichia coli.
Nucleic acids research 1998;26(7):1560-6.
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1997: Rice D W; Rafferty J B; Artymiuk P J; Lloyd R G
Insights into the mechanisms of homologous recombination from the structure of RuvA.
Current opinion in structural biology 1997;7(6):798-803.
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1997: Mahdi A A; McGlynn P; Levett S D; Lloyd R G
DNA binding and helicase domains of the Escherichia coli recombination protein RecG.
Nucleic acids research 1997;25(19):3875-80.
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1997: McGlynn P; Al-Deib A A; Liu J; Marians K J; Lloyd R G
The DNA replication protein PriA and the recombination protein RecG bind D-loops.
Journal of molecular biology 1997;270(2):212-21.
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1997: Martin F; Sharples G J; Lloyd R G; Eiler S; Moras D; Gangloff J; Eriani G
Characterization of a thermosensitive Escherichia coli aspartyl-tRNA synthetase mutant.
Journal of bacteriology 1997;179(11):3691-6.
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1997: Chan S N; Harris L; Bolt E L; Whitby M C; Lloyd R G
Sequence specificity and biochemical characterization of the RusA Holliday junction resolvase of Escherichia coli.
The Journal of biological chemistry 1997;272(23):14873-82.
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1997: Sedelnikova S E; Rafferty J B; Hargreaves D; Mahdi A A; Lloyd R G; Rice D W
Crystallization of E. coli RuvA gives insights into the symmetry of a Holliday junction/protein complex.
Acta crystallographica. Section D, Biological crystallography 1997;53(Pt 1):122-4.
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1996: Whitby M C; Bolt E L; Chan S N; Lloyd R G
Interactions between RuvA and RuvC at Holliday junctions: inhibition of junction cleavage and formation of a RuvA-RuvC-DNA complex.
Journal of molecular biology 1996;264(5):878-90.
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1996: Vincent S D; Mahdi A A; Lloyd R G
The RecG branch migration protein of Escherichia coli dissociates R-loops.
Journal of molecular biology 1996;264(4):713-21.
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1996: Al-Deib A A; Mahdi A A; Lloyd R G
Modulation of recombination and DNA repair by the RecG and PriA helicases of Escherichia coli K-12.
Journal of bacteriology 1996;178(23):6782-9.
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1996: Rafferty J B; Sedelnikova S E; Hargreaves D; Artymiuk P J; Baker P J; Sharples G J; Mahdi A A; Lloyd R G; Rice D W
Crystal structure of DNA recombination protein RuvA and a model for its binding to the Holliday junction.
Science (New York, N.Y.) 1996;274(5286):415-21.
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1996: Ryder L; Sharples G J; Lloyd R G
Recombination-dependent growth in exonuclease-depleted recBC sbcBC strains of Escherichia coli K-12.
Genetics 1996;143(3):1101-14.
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1996: Mahdi A A; Sharples G J; Mandal T N; Lloyd R G
Holliday junction resolvases encoded by homologous rusA genes in Escherichia coli K-12 and phage 82.
Journal of molecular biology 1996;257(3):561-73.
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1995: Whitby M C; Lloyd R G
Branch migration of three-strand recombination intermediates by RecG, a possible pathway for securing exchanges initiated by 3'-tailed duplex DNA.
The EMBO journal 1995;14(14):3302-10.
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1995: Lloyd R G; Buckman C
Conjugational recombination in Escherichia coli: genetic analysis of recombinant formation in Hfr x F- crosses.
Genetics 1995;139(3):1123-48.
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1995: Whitby M C; Lloyd R G
Altered SOS induction associated with mutations in recF, recO and recR.
Molecular & general genetics : MGG 1995;246(2):174-9.
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1994: Sharples G J; Chan S N; Mahdi A A; Whitby M C; Lloyd R G
Processing of intermediates in recombination and DNA repair: identification of a new endonuclease that specifically cleaves Holliday junctions.
The EMBO journal 1994;13(24):6133-42.
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1994: Whitby M C; Vincent S D; Lloyd R G
Branch migration of Holliday junctions: identification of RecG protein as a junction specific DNA helicase.
The EMBO journal 1994;13(21):5220-8.
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1994: Ryder L; Whitby M C; Lloyd R G
Mutation of recF, recJ, recO, recQ, or recR improves Hfr recombination in resolvase-deficient ruv recG strains of Escherichia coli.
Journal of bacteriology 1994;176(6):1570-7.
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1994: Sharples G J; Whitby M C; Ryder L; Lloyd R G
A mutation in helicase motif III of E. coli RecG protein abolishes branch migration of Holliday junctions.
Nucleic acids research 1994;22(3):308-13.
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1994: Dunderdale H J; Sharples G J; Lloyd R G; West S C
Cloning, overexpression, purification, and characterization of the Escherichia coli RuvC Holliday junction resolvase.
The Journal of biological chemistry 1994;269(7):5187-94.
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1993: Whitby M C; Ryder L; Lloyd R G
Reverse branch migration of Holliday junctions by RecG protein: a new mechanism for resolution of intermediates in recombination and DNA repair.
Cell 1993;75(2):341-50.
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1993: Sharples G J; Lloyd R G
An E. coli RuvC mutant defective in cleavage of synthetic Holliday junctions.
Nucleic acids research 1993;21(15):3359-64.
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1993: Mandal T N; Mahdi A A; Sharples G J; Lloyd R G
Resolution of Holliday intermediates in recombination and DNA repair: indirect suppression of ruvA, ruvB, and ruvC mutations.
Journal of bacteriology 1993;175(14):4325-34.
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1993: Sharples G J; Lloyd R G
Location of the Bacillus subtilis sbcD gene downstream of addAB, the analogues of E. coli recBC.
Nucleic acids research 1993;21(8):2010.
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1993: Lloyd R G; Sharples G J
Processing of recombination intermediates by the RecG and RuvAB proteins of Escherichia coli.
Nucleic acids research 1993;21(8):1719-25.
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1993: Lloyd R G; Sharples G J
Dissociation of synthetic Holliday junctions by E. coli RecG protein.
The EMBO journal 1993;12(1):17-22.
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1992: Lloyd R G; Sharples G J
Genetic analysis of recombination in prokaryotes.
Current opinion in genetics & development 1992;2(5):683-90.
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1992: Tsaneva I R; Illing G; Lloyd R G; West S C
Purification and properties of the RuvA and RuvB proteins of Escherichia coli.
Molecular & general genetics : MGG 1992;235(1):1-10.
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1992: Parsons C A; Tsaneva I; Lloyd R G; West S C
Interaction of Escherichia coli RuvA and RuvB proteins with synthetic Holliday junctions.
Proceedings of the National Academy of Sciences of the United States of America 1992;89(12):5452-6.
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1992: Gibson F P; Leach D R; Lloyd R G
Identification of sbcD mutations as cosuppressors of recBC that allow propagation of DNA palindromes in Escherichia coli K-12.
Journal of bacteriology 1992;174(4):1222-8.
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1992: Leach D R; Lloyd R G; Coulson A F
The SbcCD protein of Escherichia coli is related to two putative nucleases in the UvrA superfamily of nucleotide-binding proteins.
Genetica 1992;87(2):95-100.
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1991: Dunderdale H J; Benson F E; Parsons C A; Sharples G J; Lloyd R G; West S C
Formation and resolution of recombination intermediates by E. coli RecA and RuvC proteins.
Nature 1991;354(6354):506-10.
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1991: Sharples G J; Lloyd R G
Resolution of Holliday junctions in Escherichia coli: identification of the ruvC gene product as a 19-kilodalton protein.
Journal of bacteriology 1991;173(23):7711-5.
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1991: Sharples G J; Lloyd R G
Location of a mutation in the aspartyl-tRNA synthetase gene of Escherichia coli K12.
Mutation research 1991;264(3):93-6.
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1991: Lloyd R G; Sharples G J
Molecular organization and nucleotide sequence of the recG locus of Escherichia coli K-12.
Journal of bacteriology 1991;173(21):6837-43.
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1991: Lloyd R G
Linkage distortion following conjugational transfer of sbcC+ to recBC sbcBC strains of Escherichia coli.
Journal of bacteriology 1991;173(18):5694-8.
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1991: Lloyd R G
Conjugational recombination in resolvase-deficient ruvC mutants of Escherichia coli K-12 depends on recG.
Journal of bacteriology 1991;173(17):5414-8.
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1991: Connolly B; Parsons C A; Benson F E; Dunderdale H J; Sharples G J; Lloyd R G; West S C
Resolution of Holliday junctions in vitro requires the Escherichia coli ruvC gene product.
Proceedings of the National Academy of Sciences of the United States of America 1991;88(14):6063-7.
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1991: Benson F; Collier S; Lloyd R G
Evidence of abortive recombination in ruv mutants of Escherichia coli K12.
Molecular & general genetics : MGG 1991;225(2):266-72.
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1991: Lloyd R G; Buckman C
Overlapping functions of recD, recJ and recN provide evidence of three epistatic groups of genes in Escherichia coli recombination and DNA repair.
Biochimie 1991;73(2-3):313-20.
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1991: Lloyd R G; Buckman C
Genetic analysis of the recG locus of Escherichia coli K-12 and of its role in recombination and DNA repair.
Journal of bacteriology 1991;173(3):1004-11.
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1990: Sharples G J; Lloyd R G
A novel repeated DNA sequence located in the intergenic regions of bacterial chromosomes.
Nucleic acids research 1990;18(22):6503-8.
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1990: Sharples G J; Benson F E; Illing G T; Lloyd R G
Molecular and functional analysis of the ruv region of Escherichia coli K-12 reveals three genes involved in DNA repair and recombination.
Molecular & general genetics : MGG 1990;221(2):219-26.
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1989: Naom I S; Morton S J; Leach D R; Lloyd R G
Molecular organization of sbcC, a gene that affects genetic recombination and the viability of DNA palindromes in Escherichia coli K-12.
Nucleic acids research 1989;17(20):8033-45.
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1989: Mahdi A A; Lloyd R G
The recR locus of Escherichia coli K-12: molecular cloning, DNA sequencing and identification of the gene product.
Nucleic acids research 1989;17(17):6781-94.
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1989: Mahdi A A; Lloyd R G
Identification of the recR locus of Escherichia coli K-12 and analysis of its role in recombination and DNA repair.
Molecular & general genetics : MGG 1989;216(2-3):503-10.
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1988: Chalker A F; Leach D R; Lloyd R G
Escherichia coli sbcC mutants permit stable propagation of DNA replicons containing a long palindrome.
Gene 1988;71(1):201-5.
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1988: Lloyd R G; Porton M C; Buckman C
Effect of recF, recJ, recN, recO and ruv mutations on ultraviolet survival and genetic recombination in a recD strain of Escherichia coli K12.
Molecular & general genetics : MGG 1988;212(2):317-24.
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1988: Benson F E; Illing G T; Sharples G J; Lloyd R G
Nucleotide sequencing of the ruv region of Escherichia coli K-12 reveals a LexA regulated operon encoding two genes.
Nucleic acids research 1988;16(4):1541-9.
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1987: Lloyd R G; Buckman C; Benson F E
Genetic analysis of conjugational recombination in Escherichia coli K12 strains deficient in RecBCD enzyme.
Journal of general microbiology 1987;133(9):2531-8.
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1987: Lloyd R G; Evans N P; Buckman C
Formation of recombinant lacZ+ DNA in conjugational crosses with a recB mutant of Escherichia coli K12 depends on recF, recJ, and recO.
Molecular & general genetics : MGG 1987;209(1):135-41.
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1987: Rostas K; Morton S J; Picksley S M; Lloyd R G
Nucleotide sequence and LexA regulation of the Escherichia coli recN gene.
Nucleic acids research 1987;15(13):5041-9.
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1985: Lloyd R G; Buckman C
Identification and genetic analysis of sbcC mutations in commonly used recBC sbcB strains of Escherichia coli K-12.
Journal of bacteriology 1985;164(2):836-44.
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1985: Attfield P V; Benson F E; Lloyd R G
Analysis of the ruv locus of Escherichia coli K-12 and identification of the gene product.
Journal of bacteriology 1985;164(1):276-81.
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1985: Picksley S M; Morton S J; Lloyd R G
The recN locus of Escherichia coli K12: molecular analysis and identification of the gene product.
Molecular & general genetics : MGG 1985;201(2):301-7.
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1984: Lloyd R G; Benson F E; Shurvinton C E
Effect of ruv mutations on recombination and DNA repair in Escherichia coli K12.
Molecular & general genetics : MGG 1984;194(1-2):303-9.
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1984: Shurvinton C E; Lloyd R G; Benson F E; Attfield P V
Genetic analysis and molecular cloning of the Escherichia coli ruv gene.
Molecular & general genetics : MGG 1984;194(1-2):322-9.
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1984: Picksley S M; Attfield P V; Lloyd R G
Repair of DNA double-strand breaks in Escherichia coli K12 requires a functional recN product.
Molecular & general genetics : MGG 1984;195(1-2):267-74.
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1983: Lloyd R G; Picksley S M; Prescott C
Inducible expression of a gene specific to the RecF pathway for recombination in Escherichia coli K12.
Molecular & general genetics : MGG 1983;190(1):162-7.
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1982: Shurvinton C E; Lloyd R G
Damage to DNA induces expression of the ruv gene of Escherichia coli.
Molecular & general genetics : MGG 1982;185(2):352-5.
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